WOUND HEALING RATE IN ORIENTAL BEECH TREES FOLLOWING LOGGING DAMAGE

Beech is the most important commercial species in the Caspian forests of Iran. Selective cutting and harvesting methods may adversely impact the quality of the residual trees, as the injuries make the trees prone to future disease, insect infestations or timber defects. Although attempts to better understand how wounds affect the residual trees have been made in many different contexts, there are still few investigations on uneven-aged forests. In this study the key objectives were to determine and model the healing rate for different wound parameters (width, length, and area of wound); to analyse the relationship between wound healing rate (WHR), tree diameter growth and tree height growth; to analyse the WHR in relation to wound position on the stem; and to analyse the relationship between WHR, width and area of wound in DBH classes and social classes, with the aim of enabling the prognosis of logging wounds. Wounded beech trees were examined immediately after selective logging and after a 5-year period. The WHR was 31.2 +/- 7.7 cm(2) year(-1). The wound width healing rate (18.4 +/- 3.4 mm.year(-1)) was significantly higher than the wound length healing rate (4.5 +/- 1.6 mm.year(-1)). Only 12% of wounds were completely closed after a 5-year period, and 15 years are necessary for the complete closure of 80% of total wounds. The ratio of wound area to stem area at wound height (RWS) showed a more pronounced effect on diameter than on height. Regression analysis showed that WHR was correlated negatively with wound area and width and positively with tree diameter growth, but no significant relationship was found between height growth and WHR parameters. The WHR was significantly higher at an upper position than at a lower one, and statistical tests showed that the tree vertical layering classes had a significant effect on WHR. Finally, it was shown that WHRs in upper-storey trees are significantly higher than in the middle and lower storeys.