Diagnoses and discussion of the group 1 and 2 Brazilian species of Heteragrion, with descriptions of four new species (Odonata: Megapodagrionidae)

Heteragrion is the most speciose and complex genus of neotropical Megapodagrionidae, with 47 species and one subspecies, and many of them are poorly defined. To improve the knowledge of the Brazilian species of the genus, 179 specimens of 13 of the 17 described species were examined. Four new species are described in tribute to the 40th anniversary of the rock band Queen: Heteragrion freddiemercuryi (Holotype: Peruibe (24 degrees 22' 48 '' S & 47 degrees 04' 40 '' W-10 m), Sao Paulo State-09-III-2000); H. brianmayi (Holotype and allotype: Parque Estadual da Serra do Mar-Nucleo Caraguatatuba, Caraguatatuba (23 degrees 35' 36 '' S & 45 degrees 25' 07 '' W-61 m), Sao Paulo State-05-II-2000 and 10-III-2002); H. rogertaylori (Holotype and allotype: Fazenda Trabiju, Pindamonhangaba (22 degrees 50' 40 '' S & 45 degrees 31' 01 '' W-651 m), Sao Paulo State- 26-III-1999); and H. johndeaconi (Holotype and allotype: Alto Paraiso de Goias (14 degrees 10' 55 '' S & 47 degrees 38' 36 '' W-1198 m), Goias State-18-VI-2008 and 17-VI-2008), holotypes, allotypes and most paratypes deposited in FAAL. Diagnostic illustrations of all species studied are provided. Color photographs of live individuals of H. brianmayi, H. aurantiacum, H. consors, H. mantiqueirae, H. tiradentense and H. triangulare are also presented.