Revision of Bethylinae fossils (Hymenoptera: Bethylidae) from Baltic, Rovno and Oise amber, with comments on the Tertiary fauna of the subfamily

The known distribution and taxonomic variation of the previously described species Lytopsenella kerneggeri OHL are broadened. Eight new fossil species of Bethylinae are described and illustrated: Eupsenella aulax RAMOS & AZEVEDO sp. nov., E. klesoviana RAMOS & AZEVEDO sp. nov., E. rossica RAMOS & AZEVEDO sp. nov., E. yantarnica RAMOS & AZEVEDO sp. nov., Goniozus definitus RAMOS & AZEVEDO sp. nov., Lytopsenella baltica RAMOS & AZEVEDO sp. nov., L. maritima RAMOS & AZEVEDO sp. nov., and Sierola rovniana RAMOS & AZEVEDO sp. nov. Protobethylus DE PLOEG & NEL, 2004 is considered a junior synonym of Eupsenella WESTWOOD, 1874, and Protobethylus eocenicus DE PLOEG & NEL, 2004 is transferred from Protobethylus to Eupsenella as E. eocenica comb. nov. A key to the fossil species of Bethylinae is provided. This paper brings the total known fossil species of Bethylinae to 17.