A contribution to the ITS-LSU phylogeny of the genus Leucopaxillus (/tricholomatoid clade, Agaricales), with three new genera and notes on Porpoloma

Phylogenetic analyses based on ITS-LSU rDNA sequences dataset indicate that Leucopaxillus, as currently defined, is a highly polyphyletic genus. The new genera Giacomia, Notholepista and Pseudoclitopilus are introduced to accommodate Leucopaxillus mirabilis, L. subzonalis and L. rhodoleucus, respectively. Leucopaxillus subg. Aspropaxillus also seems to represent an independent evolutionary line in the /tricholomatoid clade, for which we suggest resurrecting the genus Aspropaxillus. Furthermore, the morphologically allied genus Porpoloma is also polyphyletic.