Journal
NATURE
Volume 431, Issue 7009, Pages 680-684Publisher
NATURE PUBLISHING GROUP
DOI: 10.1038/nature02855
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Tyrannosauroids are one of the last and the most successful large-bodied predatory dinosaur groups(1-5), but their early history remains poorly understood. Here we report a new basal tyrannosauroid from the Early Cretaceous Yixian Formation of western Liaoning, China, which is small and gracile and has relatively long arms with three-fingered hands. The new taxon is the earliest known unquestionable tyrannosauroid found so far(6-9). It shows a mosaic of characters, including a derived cranial structure resembling that of derived tyrannosauroids(1-5) and a primitive postcranial skeleton similar to basal coelurosaurians. One of the specimens also preserves a filamentous integumentary covering similar to that of other coelurosaurian theropods from western Liaoning. This provides the first direct fossil evidence that tyrannosauroids had protofeathers.
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