Journal
ZOOTAXA
Volume 5005, Issue 1, Pages 1-20Publisher
MAGNOLIA PRESS
DOI: 10.11646/zootaxa.5005.1.1
Keywords
Gasterosteidae; nine-spined stickleback; new species; Yamagata Prefecture; northern Japan
Categories
Funding
- JSPS KAKENHI [23570105]
- Grants-in-Aid for Scientific Research [23570105] Funding Source: KAKEN
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The new species Pungitius modestus from Japan is characterized by specific features such as the number of lateral plates, fin rays, and coloration patterns, distinguishing it from other Pungitius species.
A new species of nine-spined stickleback, Pungitius modestus, is described based on the holotype and 17 paratypes (38.7-51.7 mm standard length) collected from the inland area of Yamagata Prefecture, northern Honshu, Japan. The new species is distinguished from the other species of Pungitius by the following combination of characters: 30-32 small unconnected lateral plates; dorsal-fin rays VIII-X (usually IX)+9-11; anal-fin rays 7-10 (usually 8); pectoral-fin rays 10; a short spiny dorsal fin base (26.5%-29.8% SL); the first spiny dorsal fin spine behind the pectoral-fin base; a long pre -anal fin (59.9%-67.4% SL); the anal-fin spine below the 1st-3rd dorsal-fin rays; a short pelvic-fin spine (6.3%-9.1% SL); a short anal-fin spine (4.1%-6.0% SL); a long caudal peduncle (14.3%-19.7% SL); no body markings; membranes of the dorsal-fin spines dark brown with black pigments; the entire male body, and soft dorsal and anal fins, becoming black in the breeding season; the anteroventral process of the ectocoracoid present; and the dorsal extension of the ascending process of the pelvis level with the dorsal-most actinost.
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