A review of fossil Bursidae and their use for phylogeny calibration

Bursidae Thiele, 1925 is a moderately diverse group of extant tonnoidean gastropods with a significant fossil record. We review the fossil record of the family. We exclude some taxa from Bursidae, particularly the most ancient ones: Hanaibursa aquilana (Parona, 1909) (Aptian) and Bursa saundersi Adegoke, 1977 (Selandian). We exclude the genus Olequahia Stewart, 1926; its posterior siphonal canal is not analogous with that of Bursidae. We also discuss the possible revision of the type genus, Bursa Roding, 1798, on the basis of previously published phylogenies; the genus is not monophyletic. We create two new genera, Olssonia n. gen. (type species: Bursa chira Olsson, 1930) and Aquitanobursa n. gen. (type species: Ranella grateloupi d'Orbigny, 1852), containing only fossil species. Lectotypes are designated for Ranella grateloupi d'Orbigny, 1852, Ranella morrisi d'Archiac & Haime, 1853 and Apollon pelouatensis Cossmann & Peyrot, 1924. Based on this revision of the fossil record, we propose five fossil calibration points that can be used to date molecular phylogenetic trees of Bursidae.