The Structure, Functions, and Mechanical Properties of Keratin

Keratin is one of the most important structural proteins in nature and is widely found in the integument in vertebrates. It is classified into two types: alpha-helices and beta-pleated sheets. Keratinized materials can be considered as fiber-reinforced composites consisting of crystalline intermediate filaments embedded in an amorphous protein matrix. They have a wide variety of morphologies and properties depending on different functions. Here, we review selected keratin-based materials, such as skin, hair, wool, quill, horn, hoof, feather, and beak, focusing on the structure-mechanical property-function relationships and finally give some insights on bioinspired composite design based on keratinized materials.